01 - Preheat oven to 350°F. Grease and flour two 9-inch round cake pans, then line the bottoms with parchment paper for easy release.
02 - In a large mixing bowl, sift together the all-purpose flour, granulated sugar, unsweetened cocoa powder, baking powder, baking soda, and salt until evenly blended.
03 - Add the eggs, whole milk, vegetable oil, and vanilla extract to the dry mixture. Beat on medium speed until fully combined. Gradually pour in the boiling water while mixing continuously until the batter is smooth and uniform. The batter will be quite thin — this is expected.
04 - Divide the batter evenly between the two prepared cake pans. Bake for 30 to 35 minutes, or until a toothpick inserted into the center of each layer comes out clean. Allow the cakes to cool in their pans for 10 minutes, then turn them out onto wire racks to cool completely.
05 - While the cakes bake, toss the sliced strawberries with granulated sugar and fresh lemon juice in a bowl. Set aside for at least 20 minutes to allow the berries to release their natural juices and develop flavor.
06 - Heat the heavy cream in a saucepan over medium heat just until it begins to simmer — do not boil. Pour the hot cream over the chopped semisweet chocolate in a heatproof bowl. Let stand for 2 minutes, then stir gently until the mixture is glossy and completely smooth. Allow to cool slightly until it thickens to a pourable consistency.
07 - If using the whipped cream topping, beat the cold heavy whipping cream with powdered sugar and vanilla extract using an electric mixer until soft peaks form. Be careful not to overwhip.
08 - Place the first cake layer on a serving platter. Arrange a generous layer of the macerated strawberries over the top, reserving a few for garnish. Spread half of the chocolate ganache over the strawberries. If desired, add a layer of whipped cream on top of the strawberries before the ganache.
09 - Carefully position the second cake layer on top. Pour the remaining ganache over the top of the cake, allowing it to flow naturally down the sides. Arrange the reserved strawberries on top as garnish.
10 - Refrigerate the assembled cake for at least 1 hour to allow the layers to set and the flavors to meld. Slice into wedges and serve chilled.